Publications (1995-2019 only).

Holzwarth G, Bhandari A, Tommervik L, Macosko JC, Ornelles D, and  Lyles DS(2019). “Vesicular stomatitis virus nucleocapsids diffuse through cytoplasm by hopping from trap to trap in random directions.” Submitted, November 2, 2019

Bonin K, Smelser AM,  Moreno NS, Holzwarth G, Wang K, Levy P, and Vidi P-A(2018). Structured illumination to spatially map chromatin motions. J. Biomed. Optics23(5),056007(2018),

Yacovone SK, Smelser AM, Macosko JC, Holzwarth,G, Ornelles DA, and Lyles DS (2016). “Migration of nucleocapsids in Vesicular Stomatitis Virus-infected Cells is dependent on both Microtubules and Actin Filaments”, J. Virology 90(13),6159-6170(2016), doi:10.1128/JV1.00488-16

Martin MJ, Smelser AM, and Holzwarth G (2016). “Dividing organelle tracks into Brownian and motor-driven intervals by variational maximization of the Bayesian evidence”, Eur Biophys J 45, 269-277. DOI 10.11007/s00249-015-1091-0

Smelser AM, Macosko JC, O’Dell AP, Smyre S, Bonin K, Holzwarth G (2015). “Mechanical properties of normal versus cancerous breast cells”, Biomech Model Mechanobiol doi 10.1007/s10237-015-0677-x) (

Fallesen TL, Macosko JC and Holzwarth G. “Measuring the number and spacing of molecular motors propelling a gliding microtubule”, Phys Rev E 83.011918 (2011)(

Fallesen TL, Macosko JC and Holzwarth G. “Force-velocity relationship for multiple kinesin motors pulling a magnetic bead”, Eur Biophys J 40, 1071-1079 (2011) (

Fallesen T, Hill DB, Steen, M, Macosko JC, Bonin K, and Holzwarth G, “Magnet polepiece design for uniform magnetic force on superparamagnetic beads”, Rev Sci Instrum 81, 074303 (2010), (

Shtridelman Y, Holzwarth GM, Bauer CT, Gassman NR, DeWitt DA, and Macosko JC, “In vivo Multimotor Force-Velocity Curves by Tracking and Sizing Sub-Diffraction limited Vesicles”, Cellular and molecular Bioengineering 2(2),190-199, 2009 (

Gagliano J, Walb M, Blaker B, Macosko JC, and Holzwarth, G. “Kinesin velocity increases with the number of motors pulling against viscoelastic drag”, European Biophysics Journal 39 (5),801-813 (2010)[pdf]

Hill DB, Macosko JC, Holzwarth GM “Motion-enhanced, differential interference contrast (MEDIC) microscopy of moving vesicles in live cells: VE-DIC updated”, J. Microscopy 231, 433-439 (2008)[pdf].

Shtridelman Y, Holzwarth, GM, Bauer CT, Gassman NR, DeWitt DA, and Macosko, JC, ” In vivo Multimotor Force-Velocity Curves by Tracking and Sizing Sub-Diffraction-limited Vesicles”, Cellular and Molecular Bioengineering 2, 190-199 (2009)

Macosko, J.C., Newbern, J.M., Rockford, J., Chisena, E.N., Brown, C.M., Holzwarth, G., and Milligan C.E. “Fewer active motors per vesicle may explain slowed vesicle transport in chick motoneurons after three days in vitro”, Brain Research 1211, 6-12 (2008).

Chisena EN, Wall RA, Macosko JC, and Holzwarth GM “Speckled microtubules improve tracking in motor-protein gliding assays”, Phys Biol 4, 10-15 (2007).Holzwarth G, Bonin K, and Hill DB, “Forces Required of Kinesin during Processive Transport through Cytoplasm”, Biophys J. 82, 1784-1790(2002) [pdf].[html].

Holzwarth G, Hill DB, and McLaughlin EB, “Polarization-modulated DIC microscopy with a variable retarder”, Applied Optics 39, 6288-94(2000).

Holzwarth G, Webb SC, Kubinski DJ, and Allen NS, Improving DIC Microscopy with Polarization Modulation, J. Microscopy 188, 249-254 (1997)[pdf].

Holzwarth G, Channel thickness variations could degrade resolution in electrophoresis. Electrophoresis 17, 1587-1589(1996).

Sabanayagam CR. and Holzwarth G, Real-time velocity of DNA bands during field-inversion gel electrophoresis. Electrophoresis 17, 1052-1059 (1996).

Neitzey LM, Holzwarth G, Duke TAJ, and Viovy JL. Two-dimensional motion of DNA bands during pulsed-field gel electrophoresis: effect of field angle. Biopolymers 35, 307-317 (1995).

Hutson MS, Holzwarth G, Duke TAJ, and Viovy JL. Two-dimensional motion of DNA bands during 120-degree  pulsed-field gel electrophoresis: effect of molecular weight. Biopolymers 35, 297-306(1995).